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Table 5 Reproductive behaviors of male Iberian red deer (Cervus elaphus hispanicus; N = 72) reproductive behaviors displayed during four consecutives rutting seasons (N = 227 observations; N = 330 recorded behaviors). The number of times a behaviour was recorded is shown as Obs. (observed)

From: The dark-ventral-patch of male red deer, a sexual signal that conveys the degree of involvement in rutting behavior

 

Description

Roaring (Obs. = 146)

Loud call that males repeat frequently during the rut.

Flehmen (Obs. = 38)

The male raises his head and stretches the neck forward, and with the mouth opened the upper lip was curled back.

Antler rubbing (Obs. = 50)

Rubbing of shrubs and trees where the male sways his head sideways, sometimes picking up branches and leaves that remain in the antlers.

Parallel walk (Obs. = 10)

Two stags walk side by side usually turning to face its opponent and lowering its antlers. During the walk males are often roaring.

Fight (Obs. = 4)

Males intertwine their antlers and struggle with the opponent by shaking their heads sideways, trying to push back the rival male.

Female harassment (Obs. = 76)

The male pushes his face into the female’s anogenital region and runs after her for more than 5 s.

Mount female (Obs. = 6)

Attempts to copulate, where the male rests his body on the back of the female.